Macedoniella Drensky , 1935 with two new synonyms ( Araneae : Gnaphosidae )

Examination of the type material of the forgotten species Liocranum ochraceum Simon, 1867 reveals this species has to be transferred to the genus Anagraphis comb. nov. and that Anagraphis pallida (Hadjissarantos, 1940) is its junior synonym (syn. nov.). Furthermore, the monotypic genus Macedoniella Drensky, 1935 is a junior synonym of Anagraphis and M. karamani a junior synonym of Anagraphis ochraceum (L. Koch, 1867) (syn. nov.).

Although the arachnofauna of Europe has become better and better known, many species described in the 19 th century remain species inquirendae.These are still valid names and investigation of type material of these species must be a priority.Examination of type material in the Natural History Museum, London and a study of the papers where the species were published, provided the opportunity here to reveal the identity of the enigmatic genus Macedoniella Drensky, 1935 and its relationships to Anagraphis pallida (Hadjissarantos, 1940).
All reference material cited was collected by the author, unless indicated otherwise and is stored in his collection.Abbreviations: CRB: Collection Robert Bosmans.NHMB: Natural History Museum, London.ZMUA: Zoological Museum of the University of Athens.(Deltshev 2003).

Systematics
Holotype male, paratype female of Talanites pallidus from GREECE, Attiki, Pendeli-Dyonisos; Coll.Hadjissarantos (ZMUA); examined by Chatzaki et al. (2002).Abstract.Examination of the type material of the forgotten species Liocranum ochraceum Simon, 1867 reveals this species has to be transferred to the genus Anagraphis comb.nov.and that Anagraphis pallida (Hadjissarantos, 1940) is its junior synonym (syn.nov.).Furthermore, the monotypic genus Macedoniella Drensky, 1935 is a junior synonym of Anagraphis and M. karamani a junior synonym of Anagraphis ochraceum (L.Koch, 1867)  on of the female is without any figure and the epigyne was described as "eine hufeisenförmige Wulstung, welche vorne offen ist, in diese Öffnung ragt eine halbkreisförmige Platte herein", basically meaning: "a horseshoe-shaped chitinisation, in the anterior opening with a semi-circular plate".Probably due to the absence of figures of the epigyne, the species was never recognized or cited again.

Remarks on synonymy
Examining now the epigyne of the holotype of L. ochraceum and comparing it with my recently collected material of Anagraphis pallida (Hadjissarantos, 1940), identified with the excellent redescription of Chatzaki et al. (2002), shows that L. ochraceum has to be transferred to Anagraphis.The epigynes are identical, so there is no doubt they are the same species and Anagraphis pallida (Hadjissarantos, 1940) thus becomes a junior synonym of Anagraphis ochracea (L.Koch, 1867) comb.nov.
When examining the holotype of Liocranum ochraceum, i further came across the resemblances to the description of a species from Macedonia, Macedoniella karamani Drensky, 1935.Macedoniella is a monotypic genus and is only known from the female.According to Deltshev (2003), the type material was lost during the Second World War.Wunderlich (2011) states that the family relationships of the genus are unsure, but indicates: "Gnaphosidae?". in the World Spider Catalogue (2014) it is listed as a nomen dubium.
The description of Drensky is rather superfluous and the figures sketchy (Fig. 1).When comparing the material of Anagraphis ochracea with Drensky's description of Macedoniella karamani, size, colour, eye disposition and position of the spinnerets are identical.A special character of Macedoniella is, according to Drensky (1935), the presence of a subterminal tooth on Mt ii (plate i, a).This is not observed in the present material.in these specimens, there are 2 dorsal and 2 prolateral spines on all femora, Ti i-iii have 3 pairs of ventral spines, Mt i-iii 2 pairs of ventral spines and Ti iV and Mt iV have more additional spines.i presume that the specimen of Drensky lost the ventral spines on the Ti and Mt, except the subterminal retrolateral spine of Ti ii.The figure of the epigyne (plate 1, b) shows a horseshoe-shaped chitinisation with an anterior darkened region, almost exactly like what L. Koch wrote in the original description of Liocranum ochraceum.
The type locality of Macedoniella karamani is somewhat to the north of the newly established dis-tribution area of Anagraphis ochracea (Fig. 1), but is within a reasonable distance.The conclusion that Macedoniella karamani Drensky, 1935 is a junior synonym of Anagraphis ochracea (L.Koch, 1867) thus seems evident.

Description
i refer to the excellent redescription of Chatzaki et al. (2002).

Distribution
Anagraphis pallida appears to be a species with a relatively small distribution area (Fig. 2), but according to the number of recent localities, it is not rare.L. Koch described the female in 1867 as Liocranum ochraceum from the Greek island of Corfu.Drensky (1935) described the female again as a new species, namely Macedoniella karamani from the Republic of Macedonia.The species was then described for a third time as Talanites pallidus by Hadjissarantos (1940) from Attiki in Greece, this time from both a male and female.Chatzaki et al. (2002) redescribed it, transferred it to the genus Anagraphis and added the island Antikithyra as a new locality.The species was also cited from Albania by Deltshev et al. (2011).
Here, new records from all over the Peloponnisos and from the islands Alonissos, Evvoia, Skopelos, Kefalonia and Lefkada are added and they provide confirmation of its wide distribution in Greece.The locality in the Republic of Macedonia (FYRoM) is the northernmost site of its distribution.(1935) Liocranum ochraceum was described by L. Koch in 1867 from the Greek island of Corfu.The descripti-On the identity of the genera Anagraphis Simon, 1893 and Macedoniella Drensky